Among the key features found in primates are hands and feet that can grasp and holding objects谁能帮我分析一下这个句子的主语的结构
热心网友
这是一个倒装句，主语是hands and feet that can grasp and holding objects，其中that can grasp and holding objects是定语从句，修饰主语。
热心网友
这是一个倒装句.主语为 hands and feet 之后的从句为定语从句。Among the key feathers为介词短语做表语前置，并且the key feathers后还带着过去分词短语found in primates 对其进行修饰。所以正常语序为：Hands and feet that can grasp and holding objects are among the key features found in primates
热心网友
主语是features
热心网友
hands and feet 是主语,这是个倒装句